Glyphithyreus wetherelli is the most common crab across the London Clay formation. It is very small and can often be found in the shingle patches of the beach in small nodules. Very occasionally it can be found in large septarian nodules. It is not often preserved with claws and legs attached. It is synonyms with Plagiolophus wetherelli which is an older name for it.
It has been found at Sheppey, Butt Cliff and Maylandsea.
Description: Carapace almost circular to transversely subovate, flattened; orbitofrontal margin more than half carapace width, straight, front slightly produced with median notch; short anterolateral margins with three small spines; lobes well defined; epi- and mesobranchial lobes bar-like with posteriorly directed median spur; according to condition, bar-like metabranchial ridge continuous with cardiac region. Claws elongate, smooth.
(Description from: A guide to the fossil Decapoda (Crustacea: Axiidea, Anomura, Brachyura) of the British Isles)
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Crustacea
Superclass: Multicrustacea
Class: Malacostraca
Subclass: Eumalacostraca
Superorder: Eucarida
Order: Decapoda
Suborder: Pleocyemata
Infraorder: Brachyura
Section: Eubrachyura
Subsection: Heterotremata
Superfamily: Xanthoidea
Family: Panopeidae (Cloest living relatives)
Genus: Glyphithyreus
Species: Glyphithyreus wetherelli
Image from Gary Walker
Panopeus lacustris, a living relative of Glyphithyreus wetherelli.
Image from:
Glyphithyreus wetherelli is extinct with its closest living relatives being from the Panopeidae family often refered to as 'mud crabs'
The Panopeidae family of crabs most live in the Atlantic Ocean or eastern Pacific Ocean.
Acknowledgments:
Thank you very much Gary Walker for allowing usage of the images on this page.
References:
https://www.gbif.org/occurrence/417138481
https://www.mindat.org/taxon-4646648.html
[A monograph of the fossil malacostracous Crustacea of Great Britain]
https://science.mnhn.fr/institution/mnhn/collection/f/item/r11524?listIndex=24&listCount=31
https://marinespecies.org/aphia.php?p=taxdetails&id=106765
https://en.wikipedia.org/wiki/Panopeidae
https://www.inaturalist.org/photos/7873464
https://www.sciencedirect.com/science/article/pii/S0016787819301191#fig0020
https://marinespecies.org/aphia.php?p=taxdetails&id=1619313
Further reading:
https://www.mindat.org/taxon-4646647.html
Papers:
Joe S.H. Collins, Claire J.T. Mellish, Andrew J. Ross, Phillip R. Crabb, Stephen K. Donovan (2020), A guide to the fossil Decapoda (Crustacea: Axiidea, Anomura, Brachyura) of the British Isles
M'Coy (1849), British fossil Crustacea